(An Illustrated Key To the Philippine Freshwater Zooplankton.)

Back to key





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+Aquino, M. R. Y, Cho, C. D., Cruz, M. A. S., Saquiquit, A. G. and Papa, R. D. S. 2008b. Zooplankton Composition and Diversity in Paoay Lake, Luzon Is., Philippines.
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+Brehm, V. 1933c. Die Cladoceran der Deutschen Limnologischen Sunda Expedition.
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+Brehm, V. 1933d. Mitteilungen der Wallacea-Expedition Woltereck. Mitteilung VI: Die Alona- und Alonella-Arten von Dagiangan.
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+Brehm, V. 1934b. Mitteilungen der Wallacea-Expedition Woltereck.
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Briones, J. C., C.-H. Tsai, T. Nakazawa, Y. Sakai, R. D. S. Papa, C.-h. Hsieh, N. Okuda. 2012.
Long-Term Changes in the Diet of Gymnogobius isaza from Lake Biwa, Japan: Effects of Body Size and Environmental Prey Availability
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Esquivel, A., A. Barani, M. Macek, R. Soto-Castor, C. Bulit. 2016.
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Fernando. C. H. 1980. The Freshwater Zooplankton of Sri Lanka, with Discussion of Tropical Freshwater Zooplankton Composition.
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Flössner, D. 1972. Krebstiere, Crustacea, Kiemen- ­und Blattfusser, Branchiopoda, Fishlause, Brachiura.
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Frey, D. G. 1982. Relocation of Chydorus barroisi and related species, (Cladocera, Chydoridae) to a new genus and description of two new species.
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Korovchinsky, Nikolai M. 2013. Cladocera (Crustacea: Branchiopoda) of South East Asia: history of exploration, taxon richness and notes on zoogeography.

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Lai, H.C. and Fernando, C.H. 1979. The Malaysian Tropodiaptomus (Copepoda: Calanoida) and its distribution.
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+Legaspi, K., A. Y. A. Lau, P. Jordan, A. Mackay4, S. Mcgowan, G. Mcglynn, S. Baldia, R. D. Papa and D. Taylor. 2015b.
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Leon, J. R. de, H. G. de Vera, E. J. A. Giron, H. J. A. Guerrero,
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Temporal Variability of Abundance, Morphological and
Reproductive Traits of the Invasive Arctodiaptomus dorsalis
(Marsh 1907) (Copepoda: Calanoida: Diaptomidae) in Relation to
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Nielsen, B. H., Santiago A. E. and Petersen F. 1981. The Hydraulic Controle Structure. A Threat to the Fishpen Industry in Laguna de Bay.
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Nielsen, B.H., 1981 Laguna de Bay is born again.
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Nogrady, T and Segers H. 2002. Rotifera.
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  Continental Waters of the World vol. 18. 264




+Orlova-Bienkowskaja MY, 2001f. Cladocera: Anomopoda Daphniidae:
genus Simocephalus.
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Pagano, M., Saint-Jean, L., Arfi, A., Bouvy, M.  & Guiral, D. 1999. Zooplankton food limitation and grazing impact in a eutrophic
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Feeding of tropical cladocerans (Moina micrura, Diaphanosoma excisum) and rotifer (Brachionus calyciflorus) on natural phytoplankton: effect of phytoplankton size–structure.
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+Palomar, N. E. and Mamaril, A. C.  2001e. Autoecology of an endemic calanoid copepod: the first step in biodiversity conservation in lakes. In CB Santiago, ML Cuvin-Aralar and ZU Basiao (Eds.). Conservation and Ecological Management of Philippine Lakes in Relation to Fisheries and Aquaculture (p. 155). Southeast Asian Fisheries Development Center, Aquaculture Department, Iloilo, Philippines; Philippine Council for Aquatic and Marine Research and Development, Los Baños, Laguna, Philippines; and Bureau of Fisheries and Aquatic Resources, Quezon City, Philippines. Review: (Visited December 2014)


+Panogadia-Reyes C. M., Cruz, E. I. and Bautista, S. L. 2004a,
Philippine Species of Mesocyclops (Crustacea: Copepoda) as a Biological Control Agent of Aedes aegypti (Linnaeus) Dengue
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+Papa, R. D. S. 2005a. A Study on the Zooplankton Diet of the Freshwater Sardine, Sardinella tawilis (Herre 1927) in Lake Taal, Batangas.  University of Santo Tomas España. MSc thesis


+Papa, R. D., Wu, J. T., Baldia, S. F. Cho, C., Cruz, M. A.
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Papa, R. D., Pagulayan, R., and Pagulayan, A. E. 2007. Abundance and Composition of Zooplankton (Copepoda, Cladocera, Rotifera) in Lake Taal: Potential Impact of Intensive Size-Selective Predation by the Freshwater Sardine, Sardinella tawilis (Herre 1927).
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+Papa, R. D., Pagulayan, R. C., and Pagulayan, A. E. J., 2008a
Zooplanktivory in the Endemic Freshwater Sardine, Sardinella tawilis (Herre 1927) of Taal Lake, the Philippines.
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+Papa, Rey Donne S. and  Mamaril, Augustus C., 2008c.
Methods in Zooplankton Sampling, Ecology and Identification for General Biology Teachers.
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+Papa, R. D., Wu, Jiunn-Tzong, Baldia S., Cho, C., Cruz, M. A.,
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Blooms of the Colonial Green Algae, Botryococcus braunii Kützing, in Paoay Lake, Luzon Island, Philippines.
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+Papa, R. S. D. and A. C. Mamaril Sr. 2011a.
History of the biodiversity and limno-ecological studies on Lake Taal with notes on the current state of Philippine limnology.
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+Papa, R. S. D., M. T. Zafaralla and R. Eckmann, 2011b.
Spatio-temporal variation of zooplankton community in a tropical caldera lake with intensive aquaculture (Lake Taal, Philippines)
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+Papa, R. S. D. and M. T. Zafaralla 2011c
The composition, diversity and community dynamics of limnetic zooplankton in a tropical caldera lake (lake Taal, Philippines)
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+Papa, R. D. S., D. T. Tordesilla, L. D. Vitug, J. C. A. Briones, M. T. Zafaralla, and R. Kada. 2012a.
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+Papa, R. D. S., Dino T. Tordesillas and Augustus C. Mamaril, Sr. 2012b
An Updated Taxonomic Account of Limnetic Crustacean Zooplankton in Lake Taal, Philippines. Philippine Journal of Science 141(2):243-252



+Papa, R. D.S., H. Li, D. T. Tordesillas, Boping Han, H. J. Dumont. 2012c.
Massive Invasion of Arctodiaptomus dorsalis (Copepoda, Calanoida, Diaptomidae) in Philippine Lakes:
A Threat to Asian Zooplankton Biodiversity?
Biol Invasions 14:2471-2478
[Mistaken named 2012a before this correction - 4th November 2015]



+Papa, R. D. S., M. K. Hołyńska. 2013.
An overview of the limnetic Cyclopidae (Crustacea, Copepoda) of the Philippines, with emphasis on Mesocyclops.
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Papa, R. D. S. & J. C. A. Briones, 2014c.
Climate and human-induced changes to lake ecosystems: what we can learn from monitoring zooplankton ecology.
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+Papa, R. D. S.,K. L. Legaspi, C. G. Blaza, F. S. Lirio, N. G. P.Bangalan & T. C. Walter4. 2017a.
The abundance and morphometric characteristics of the endemic Pseudodiaptomus brehmi Kiefer, 1938 (Copepoda: Calanoida: Pseudodiaptomidae) in Lake Taal (Luzon Is., Philippines)
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+Pascual, Jhaydee Ann F., Rizo,Eric Zeus C.,Han, Boping, DumontHenri J.,& Papa, Rey Donne S. 2014a. Taxonomy and distribution of four Cladoceran families (Branchiopoda:Cladocera: Moinidae, Bosminidae, Chydoridae and Sididae) in Philippine inland waters. RAFFLES BULLETIN OF ZOOLOGY 62: 771–794
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+Paz, E. S. P. Dela, M. K. Hołynska and R. D. S. Papa. 2016b.
Mesocyclops and Thermocyclops (Copepoda, Cyclopidae)
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Peard, T. 2007. Freshwater jellyfish:  Welcome to our freshwater jellyfish (Craspedacusta sowerbii) web page
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+Petersen, F. 1980a. An illustrated list of Philippine Freshwater Zooplankton. SEAFDEC, BINANGONAN. Annual Report 1980, Annex.
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Petersen, Flemming 1980b
Kinesiske karpedamme.
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Petersen, F. 1981e Zooplankton Studies. 1980-81
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Petersen, F. 1983a. Population dynamics and production of Daphnia galeata (Crustacea, Cladocera) in Lake Esrom.
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Petersen, Flemming 1983b
Manipulerede blåmuslinger - en kommentar.
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+Petersen, F. and Carlos, M. 1984a. A Review of Zooplankton in the Philippine Lakes.
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+Petersen, F. 2007b(2014, 9th ed.). An Illustrated Key to the Philippine Freshwater Zooplankton. Including some brackish water species from Laguna de Bay. With ecological notes. [This publication]. Web-edition:
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Reddy Y. R. and Radhakrishna, Y. 1982. Redescription and/or remarks on four species of Pseudodiaptomus Herrick (Copepoda: Calanoida) from South India
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Reisinger, E. 1972. Süsswassermedusen. In: Das Zooplankton der Binnengewasser. 1. Teil.
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Richter, H. 2001. Seasonal variation in growth, quantitative and qualitative food consumption of milkfish, Chanos chanos (Forsskål 1775), and Nile tilapia, Oreochromis niloticus (L. 1758), in Laguna de Bay, Philippines
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+Rizo, E. Z. C., J. A. F. Pascual,.D. J. B. Dy, L. J. A. Labicane,.C. B. Trinidad, and R. D. S. Papa. 2015a.
New records of an Invasive Calanoid Copepod, Arctodiaptomus dorsalis (Marsh, 1907) in freshwater ecosystems in the Bicol Peninsula (Luzon Is., Philippines)
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+Rizo, E. Z., Y. Gu, R. D. S. Papa, H. J. Dumont, B.-P. Han. 2017b
Identifying functional groups and ecological roles of tropical and subtropical freshwater Cladocera in Asia
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Ruttner-Kolisko, A. 1972. Rotatoria. In: Das Zooplankton der Binnengewasser.
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Santiago, C. B., Gonzal, A. C., Aralar, E. V.  & Arcilla, R. P. 2004.
Effect of stunting of juvenile bighead carp
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Sarma, S. S. S. 1988. New records of freshwater rotifers (Rotifera) from Indian waters
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+Schiemer, F., D. Simon, U.S. Amarasinghe & J. Moreau. (eds.) 2008e.
Aquatic ecosystems and development: comparative Asian perspectives.
Chapter 8: Jacobus Vijverberg, P. Bandu Amarasinghe, Tanaporn Chitapalapong, Roberto C. Pagulayan, M. Gamage Ariyaratne, Edwin Rey J. Pamanian, E. Ivan L. Silva & Leo A.J. Nagelkerke: Structure of microcrustacean zooplankton communities in five tropical Asian water bodies 153.(Refered in 2011c) (Not yet red)
Chapter 9: P. Bandu Amarasinghe, M. Gamage Ariyaratne, Tanaporn Chitapalapong & Jacobus Vijverberg: Production, biomass and productivity of copepods and cladocerans in tropical Asian water bodies and the carrying capacity for zooplanktivorous fish
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(Not yet red)


+Schoenly, K.G., H.D. Justo, Jr., A.T. Barrion, M.K. Harris & D.G. Bottrell 1998b. Analysis of invertebrate biodiversity in a Philippine farmer's irrigated rice field.
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Segers, H. 2002a. The nomenclature of the Rotifera: annotated checklist of valid family and genus-group names
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December 2014)


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Effect of temperature on life history traits of the invasive calanoid copepod Arctodiaptomus dorsalis (Marsh, 1907)from Lake Taal, Philippines.
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UST Research Center for the Natural and Applied Sciences, University of Santo Tomas 2013.
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Victor, R., 1979. The taxonomy and distribution of freshwater Ostracoda (Crustacea, Ostracoda) of Malaysia, Indonesia and the Philippines.
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Wallace, R. L., Snell, T. W., Ricci, C. & Nogrady, T.
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Copepod web portal: (Visited December 2014)


Krepsdyr I ferskvann (Crustacean In Freshwater):

(Visited December 2014)



The world of copepod:  (Visited December 2014)

 WoRMS (


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